J Immunol 2005,
PMID: 15843522
Kawauchi, Kiyotaka; Ihjima, Kimiko; Yamada, Osamu
Human telomerase activity is induced by Ag receptor ligation in T and B cells. However, it is unknown whether telomerase activity is increased in association with activation and proliferation of NK cells. We found that telomerase activity in a human NK cell line (NK-92), which requires IL-2 for proliferation, was increased within 24 h after stimulation with IL-2. Levels of human telomerase reverse transcriptase (hTERT) mRNA and protein correlated with telomerase activity. ERK1/2 and Akt kinase (Akt) were activated by IL-2 stimulation. LY294002, an inhibitor of PI3K, abolished expression of hTERT mRNA and protein expression and abolished hTERT activity, whereas PD98059, which inhibits MEK1/2 and thus ERK1/2, had no effect. In addition, radicicol, an inhibitor of heat shock protein 90 (Hsp90), and rapamycin, an inhibitor of the mammalian target of rapamycin (mTOR), blocked IL-2-induced hTERT activity and nuclear translocation of hTERT but not hTERT mRNA expression. hTERT was coimmunoprecipitated with Akt, Hsp90, mTOR, and p70 S6 kinase (S6K), suggesting that these molecules form a physical complex. Immunoprecipitates of Akt, Hsp90, mTOR, and S6K from IL-2-stimulated NK-92 cells contained telomerase activity. Furthermore, the findings that Hsp90 and mTOR immunoprecipitates from primary samples contained telomerase activity are consistent with the results from NK-92 cells. These results indicate that IL-2 stimulation induces hTERT activation and that the mechanism of IL-2-induced hTERT activation involves transcriptional or posttranslational regulation through the pathway including PI3K/Akt, Hsp90, mTOR, and S6K in NK cells.
Diseases/Pathways annotated by Medline MESH: Leukemia, Lymphocytic, Chronic, B-Cell, Leukemia, Lymphoid
Document information provided by NCBI PubMed
Text Mining Data
telomerase reverse transcriptase → IL-2: "
IL-2 increases human
telomerase reverse transcriptase activity transcriptionally and posttranslationally through phosphatidylinositol 3'-kinase/Akt, heat shock protein 90, and mammalian target of rapamycin in transformed NK cells
"
3'-kinase/Akt → IL-2: "
IL-2 increases human telomerase reverse transcriptase activity transcriptionally and posttranslationally through phosphatidylinositol 3'-kinase/Akt , heat shock protein 90, and mammalian target of rapamycin in transformed NK cells
"
mammalian target of rapamycin → IL-2: "
IL-2 increases human telomerase reverse transcriptase activity transcriptionally and posttranslationally through phosphatidylinositol 3'-kinase/Akt, heat shock protein 90, and mammalian target of rapamycin in transformed NK cells
"
ERK1/2 → IL-2: "
ERK1/2 and Akt kinase ( Akt ) were activated by IL-2 stimulation
"
Akt kinase ( Akt ) → IL-2: "
ERK1/2 and Akt kinase ( Akt ) were activated by IL-2 stimulation
"
Manually curated Databases
-
IRef Biogrid Interaction:
TERT
—
HSP90AA1
(physical association, affinity chromatography technology)
-
IRef Biogrid Interaction:
AKT1
—
HSP90AA1
(physical association, affinity chromatography technology)
-
IRef Biogrid Interaction:
TERT
—
MTOR
(physical association, affinity chromatography technology)
-
IRef Biogrid Interaction:
AKT1
—
TERT
(physical association, affinity chromatography technology)
-
IRef Biogrid Interaction:
TERT
—
RPS6KB1
(physical association, affinity chromatography technology)
-
NCI Pathway Database Regulation of Telomerase:
Erk1-2 (MAPK3/MAPK1)
→
Erk1-2-active (MAPK3/MAPK1)
(modification, collaborate)
Evidence: assay, physical interaction
-
NCI Pathway Database Regulation of Telomerase:
Erk1-2 (MAPK3/MAPK1)
→
IL2 (IL2)
(modification, collaborate)
Evidence: assay, physical interaction
-
NCI Pathway Database IL2 signaling events mediated by PI3K:
TERT (TERT)
→
Hsp90 (HSP90AA1)
(modification, collaborate)
Evidence: mutant phenotype, assay, physical interaction
-
NCI Pathway Database IL2 signaling events mediated by PI3K:
TERT (TERT)
→
AKT1 (AKT1)
(modification, collaborate)
Evidence: mutant phenotype, assay, physical interaction
-
NCI Pathway Database IL2 signaling events mediated by PI3K:
TERT (TERT)
→
AKT1/mTOR/p70S6K/Hsp90/TERT complex (AKT1-MTOR-RPS6KB1-HSP90AA1-TERT)
(modification, collaborate)
Evidence: mutant phenotype, assay, physical interaction
-
NCI Pathway Database IL2 signaling events mediated by PI3K:
TERT (TERT)
→
mTOR (MTOR)
(modification, collaborate)
Evidence: mutant phenotype, assay, physical interaction
-
NCI Pathway Database IL2 signaling events mediated by PI3K:
TERT (TERT)
→
p70S6K (RPS6KB1)
(modification, collaborate)
Evidence: mutant phenotype, assay, physical interaction
-
NCI Pathway Database IL2 signaling events mediated by PI3K:
Hsp90 (HSP90AA1)
→
AKT1 (AKT1)
(modification, collaborate)
Evidence: mutant phenotype, assay, physical interaction
-
NCI Pathway Database IL2 signaling events mediated by PI3K:
Hsp90 (HSP90AA1)
→
AKT1/mTOR/p70S6K/Hsp90/TERT complex (AKT1-MTOR-RPS6KB1-HSP90AA1-TERT)
(modification, collaborate)
Evidence: mutant phenotype, assay, physical interaction
-
NCI Pathway Database IL2 signaling events mediated by PI3K:
Hsp90 (HSP90AA1)
→
mTOR (MTOR)
(modification, collaborate)
Evidence: mutant phenotype, assay, physical interaction
-
NCI Pathway Database IL2 signaling events mediated by PI3K:
Hsp90 (HSP90AA1)
→
p70S6K (RPS6KB1)
(modification, collaborate)
Evidence: mutant phenotype, assay, physical interaction
-
NCI Pathway Database IL2 signaling events mediated by PI3K:
AKT1 (AKT1)
→
AKT1/mTOR/p70S6K/Hsp90/TERT complex (AKT1-MTOR-RPS6KB1-HSP90AA1-TERT)
(modification, collaborate)
Evidence: mutant phenotype, assay, physical interaction
-
NCI Pathway Database IL2 signaling events mediated by PI3K:
AKT1 (AKT1)
→
mTOR (MTOR)
(modification, collaborate)
Evidence: mutant phenotype, assay, physical interaction
-
NCI Pathway Database IL2 signaling events mediated by PI3K:
AKT1 (AKT1)
→
p70S6K (RPS6KB1)
(modification, collaborate)
Evidence: mutant phenotype, assay, physical interaction
-
NCI Pathway Database IL2 signaling events mediated by PI3K:
AKT1/mTOR/p70S6K/Hsp90/TERT complex (AKT1-MTOR-RPS6KB1-HSP90AA1-TERT)
→
mTOR (MTOR)
(modification, collaborate)
Evidence: mutant phenotype, assay, physical interaction
-
NCI Pathway Database IL2 signaling events mediated by PI3K:
AKT1/mTOR/p70S6K/Hsp90/TERT complex (AKT1-MTOR-RPS6KB1-HSP90AA1-TERT)
→
p70S6K (RPS6KB1)
(modification, collaborate)
Evidence: mutant phenotype, assay, physical interaction
-
NCI Pathway Database IL2 signaling events mediated by PI3K:
mTOR (MTOR)
→
p70S6K (RPS6KB1)
(modification, collaborate)
Evidence: mutant phenotype, assay, physical interaction
-
NCI Pathway Database Regulation of Telomerase:
IL2 (IL2)
→
AKT1 (AKT1)
(modification, activates)
Evidence: assay
In total, 21 gene pairs are associated to this article in curated databases